References

[1]PARK T,SEO S,SHIN T,et al.Molecular cloning,purification,expression,and characterization ofβ-1,4-endoglucanase gene(Cel5A)fromEubacterium cellulosolvens sp.isolated from Holstein steers′rumen[J].Asian-Australasian Journal of Animal Sciences,2018,31(4):607-615．

[2]SHETTY K,PALIYATH G,POMETTO A,et al.Food Biotechnology[M].Taylor&Francis Group,LLC,2006．

[3]李燕红，赵辅昆．纤维素酶的研究进展[J]．生命科学，2005,17(5):392-397.LI Y H,ZHAO F K.Advances in cellulase research[J].Chinese Bulletin of Life Sciences,2005,17(5):392-397．

[4]陈洪章．纤维素生物技术---理论与实践[M]．北京：化学工业出版社，2014．

[5]MANSFIELD S D,MEDER R.Cellulose hydrolysis-the role of monocomponent cellulases in crystalline cellulose degradation[J].Cellulose,2003,10(2):159-169．

[6]FLEMING K,GRAY D G,MATTHEWS S.Cellulose crystallites[J].Chemistry-A European Journal,2001,7(9):1831-1835．

[7]MAZEAU K,HEUX L.Molecular dynamics simulations of bulk native crystalline and amorphous structures of cellulose[J].The Journal of Physical Chemistry B,2003,107(10):2394-2403．

[8]AGBOR V B,CICEK N,SPARLING R,et al.Biomass pretreatment:fundamentals toward application[J].Biotechnology Advances,2011,29:675-685．

[9]SINGH A,BAJAR S,DEVI A,et al.An overview on the recent developments in fungal cellulase production and their industrial applications[J].Bioresource Technology Reports,2021,14:100652．

[10]高祥照，马文奇，马常宝，等．中国作物秸秆资源利用现状分析[J]．华中农业大学学报，2002,21(3):242-247.GAO X Z,MA W Q,MA C B,et al.Analysis on the current status of utilization of crop straw in China[J].Journal of Huazhong Agricultural University,2002,21(3):242-247．

[11]KARIMI K,KHERADMANDINIA S,TAHERZADEH M J.Conversion of rice straw to sugars by dilute-acid hydrolysis[J].Biomass and Bioenergy,2006,30(3):247-253．

[12]陈庆今，刘焕彬，胡勇有．瘤胃微生物对纤维素降解机理[J]．微生物学杂志，2002,22(2):44-46.CHEN Q J,LIU H B,HU Y Y.Cellulose degradation mechanism of rumen microbes[J].Journal of Microbiology,2002,22(2):44-46．

[13]陈展．秸秆堆肥中纤维素降解菌的筛选及组合[D]．北京：中国农业大学，2005.CHEN Z.Screening and mixed culturing of cellulose decomposition microbes in the straw composting[D].Beijing:China Agricultural University,2005．

[14]HUANG C,ZONG M H,WU H,et al.Microbial oil production from rice straw hydrolysate by Trichosporon fermentans[J].Bioresource Technology,2009,100(19):4535-4538．

[15]REESE E T.Polysaccharases and the Hydrolysis of Insoluble Substrates[M].Biological Transformation of Wood by Microorganisms.Springer,Berlin,Heidelberg,1975:165-181．

[16]ENARI T M,NIKU-PAAVOLA M L.Enzymatic hydrolysis of cellulose:is the current theory of the mechanisms of hydrolysis valid?[J].Critical Reviews in Biotechnology,1987,5(1):67-87．

[17]ZHANG Y H P,LYND L R.Toward an aggregated understanding of enzymatic hydrolysis of cellulose:noncomplexed cellulase systems[J].Biotechnology and Bioengineering,2004,88(7):797-824．

[18]JUTURU V,WU J C.Microbial cellulases:engineering,production and applications[J].Renewable and Sustainable Energy Reviews,2014,33:188-203．

[19]LYND L R,WEIMER P J,van ZYL W H,et al.Microbial cellulose utilization:fundamentals and biotechnology[J].Microbiology and Molecular Biology Reviews,2002,66(3):506-577．

[20]任红梅，刘左军，袁惠君，等．低温纤维素降解菌的筛选及其产酶研究[J]．食品工业科技，2013,34(19):127-130,134.REN H M,LIU Z J,YUAN H J,et al.Study on screening and cellulase production of cold-adaped bacterium[J].Science and Technology of Food Industry,2013,34(19):127-130,134．

[21]TRUJILLO M E,FERNANDEZ-MOLINERO C,VELAZQUEZE,et al.Micromonospora mirobrigensis sp.nov.[J].International Journal of Systematic and Evolutionary Microbiology,2005,55(2):877-880．

[22]薛林贵，杨蕊琪，马高高，等．秸秆的生物降解机理及其功能微生物菌群研究进展[J]．生态科学，2017,36(3):193-199.XUE L G,YANG R Q,MA G G,et al.Progress in research of straw biodegradation mechanisms and functional microbial flora[J].Ecological Science,2017,36(3):193-199．

[23]赵旭，王文丽，李娟，等．低温秸秆降解微生物菌剂的研究进展[J]．生物技术通报，2014(11):55-61.ZHAO X,WANG W L,LI J,et al.Advanced in microbiological agent of straw degradation under low temperature[J].Biotechnology Bulletin,2014(11):55-61．

[24]王珊珊，金昱言，赵凯．高效玉米秸秆降解复合菌系的筛选[J]．黑龙江农业科学，2016(4):39-41.WANG S S,JIN Y Y,ZHAO K.Screening of composite microbial system for efficient degradation of maize straw[J].Heilongjiang Agricultural Sciences,2016(4):39-41．

[25]李燕红．福寿螺（Ampullaria crossean）内切-β-1,4-葡聚糖酶的研究[D]．北京：中国科学院上海生命科学研究院，2005.LI Y H.The studies on the endo-β-1,4-glucanase from the mollusca Ampullaria crosseans[D].Beijing:Shanghai Institutes for Biological Sciences,Chinese Academy of Sciences,2005．

[26]JUTURU V,WU J C.Microbial cellulases:engineering,production and applications[J].Renewable and Sustainable Energy Reviews,2014,33:188-203．

[27]BARR B K,HSIEH Y L,GANEM B,et al.Identification of two functionally different classes of exocellulases[J].Biochemistry,1996,35(2):586-592．

[28]KIPPER K,V?LJAM?E P,JOHANSSON G.Processive action of cellobiohydrolase Cel7Afrom Trichoderma reesei is revealed as′burst′kinetics on fluorescent polymeric model substrates[J].Biochemical Journal,2005,385(Pt2):527-535．

[29]ROUVINEN J,BERGFORS T,TEERI T,et al.Three-dimensional structure of cellobiohydrolase-ⅡfromTrichoderma reesei[J].Science,1990,249(4967):380-386．

[30]HENRIKSSON G,NUTT A,HEMRIKSSON H,et al.Endoglucanase 28(Cel12A),a new Phanerochaete chrysosporium cellulase[J].European Journal of Biochemistry,1999,259(1/2):88-95．

[31]WANG M Y,LIU K,DAI L,et al.The structural and biochemical basis for cellulose biodegradation[J].Journal of Chemical Technology&Biotechnology,2013,88(4):491-500．

[32]孙毅，金湘，毛培宏．细菌纤维素酶的结构和功能及菌种改良研究[J]．生物技术，2005,15(3):87-89.SUN Y,JIN X,MAO P H.Studies on the structures and functions of the cellulase from bacteria and the strain improvement[J].Biotechnology,2005,15(3):87-89．

[33]舒月力．纤维素酶在毕赤酵母中的异源表达及其协同作用的研究[D]．天津：天津科技大学，2019.SHU Y L.Heterologous expression of cellulase in Pichia pastoris yeast and synergistic effect[D].Tianjin:Tianjin University of Science and Technology,2019．

[34]MESSNER R,KUBICEK-PRANZ E M,GSUR A,et al.CellobiohydrolaseⅡis the main conidial-bound cellulase in Trichoderma reesei and other Trichoderma strains[J].Archives of Microbiology,1991,155(6):601-606．

[35]宋贤良，温其标，朱江．纤维素酶法水解的研究进展[J]．河南工业大学学报（自然科学版），2001,22(4):67-71,84.SONG X L,WEN Q B,ZHU J.Study progress of enzymic hydrolysis for cellulose[J].Journal of Henan University of Technology(Natural Science Edition),2001,22(4):67-71,84．

[36]DIVNE C,ST?HLBERG J,REINIKAINEN T,et al.The threedimensional crystal structure of the catalytic core of cellobiohydrolaseⅠfrom Trichoderma reesei[J].Science,1994,265(5171):524-528．

[37]VITORINO L C,BESSA L A.Microbial diversity:the gap between the estimated and the known[J].Diversity,2018,10(2):46．

[38]WEILAND-BR?UER N.Friends or foes-microbial interactions in nature[J].Biology,2021,10(6):496．

[39]LI F,XIE Y J,GAO X,et al.Screening of cellulose degradation bacteria from Min pigs and optimization of its cellulase production[J].Electronic Journal of Biotechnology,2020,48:29-35．

[40]SAHA S,ROY R N,SEN S K,et al.Characterization of cellulase-producing bacteria from the digestive tract of tilapia,Oreochromis mossambica(Peters)and grass carp,Ctenopharyngodon idella(Valenciennes)[J].Aquaculture Research,2006,37(4):380-388．

[41]SREENA C P,RESNA N K,SEBASTIAN D.Isolation and characterization of cellulase producing bacteria from the gut of termites(Odontotermes and Heterotermes species)[J].British Biotechnology Journal,2015,9(1):1-10．

[42]HOSSAIN M A,AHAMMED M A,SOBUJ S I,et al.Cellulase producing bacteria isolation,screening and media optimization from local soil sample[J].American Journal of Microbiological Research,2021,9(3):62-74．

[43]RATHORE S S,MANNIVANNAN A,NARENDHIRAKANNANR T.Screening of cellulase producing microorganisms from lake area containing water hyacinth for enzymatic hydrolysis of cellulose[J].Journal of Advanced Scientific Research,2014,5(3):23-30．

[44]PRIYA I,DHAR M K,BAJAJ B K,et al.Cellulolytic activity of thermophilic bacilli isolated from Tattapani hot spring sediment in North West Himalayas[J].Indian Journal of Microbiology,2016,56(2):228-231．

[45]CASSELLS A C,DOYLE B M.Genetic engineering and mutation breeding for tolerance to abiotic and biotic stresses:science,technology and safety[C]//Proceeding of the European Workshop on Environmental Stress and Sustainable Agriculture,Bulgaria,2002:52-82．

[46]VU V H,PHAM T A,KIM K.Improvement of fungal cellulase production by mutation and optimization of solid state fermentation[J].Mycobiology,2011,39(1):20-25．

[47]RAGHUWANSHI S,DESWAL D,KARP M,et al.Bioprocessing of enhanced cellulase production from a mutant of Trichoderma asperellum RCK2011and its application in hydrolysis of cellulose[J].Fuel,2014,124:183-189．

[48]LI X H,YANG H J,ROY B,et al.Enhanced cellulase production of the Trichoderma viride mutated by microwave and ultraviolet[J].Microbiological Research,2010,165(3):190-198．

[49]周利斌．重离子束辐照对植物组织培养及转基因操作的影响[D]．北京：中国科学院近代物理研究所，2006.ZHOU L B.Effects of heavy ion beam irradiation on plant tissue culture and gene transfer operation[D].Beijing:Institute of Modern Physics,Chinese Academy of Sciences,2006．

[50]CAO G Z,ZHANG M M,MIAO J S,et al.Effects of X-ray and carbon ion beam irradiation on membrane permeability and integrity in Saccharomyces cerevisiae cells[J].Journal of Radiation Research,2015,56(2):294-304．

[51]DONG M Y,WANG S Y,XIAO G Q,et al.Cellulase production by Aspergillus fumigatus MS13.1 mutant generated by heavy ion mutagenesis and its efficient saccharification of pretreated sweet sorghum straw[J].Process Biochemistry,2019,84:22-29．

[52]WANG S Y,JIANG B L,ZHOU X,et al.Study of a high-yield cellulase system created by heavy-ion irradiation-induced mutagenesis of Aspergillus niger and mixed fermentation with Trichoderma reesei[J].PLoS One,2015,10(12):e0144233．

[53]GADGIL N J,DAGINAWALA H F,CHAKRABARTI T,et al.Enhanced cellulase production by a mutant of Trichoderma reesei[J].Enzyme&Microbial Technology,1995,17(10):942-946．

[54]JIANG X D,GENG A L,HE N,et al.New isolate of Trichoderma viride strain for enhanced cellulolytic enzyme complex production[J].Journal of Bioscience and Bioengineering,2011,111(2):121-127．

[55]FANG H,XIA L M.High activity cellulase production by recombinant Trichoderma reesei ZU-02 with the enhanced cellobiohydrolase production[J].Bioresource Technology,2013,144:693-697．

[56]MA L,ZHANG J,ZOU G,et al.Improvement of cellulase activity in Trichoderma reesei by heterologous expression of a betaglucosidase gene from Penicillium decumbens[J].Enzyme and Microbial Technology,2011,49(4):366-371．

[57]WANG B B,XIA L M.High efficient expression of cellobiase gene fromAspergillus niger in the cells of Trichoderma reesei[J].Bioresource Technology,2011,102(6):4568-4572．

[58]SUTO M,TOMITA F.Induction and catabolite repression mechanisms of cellulase in fungi[J].Journal of Bioscience and Bioengineering,2001,92(4):305-311．

[59]DONG M Y,WANG S Y,XU F Q,et al.Efficient utilization of waste paper as an inductive feedstock for simultaneous production of cellulase and xylanase by Trichoderma longiflorum[J].Journal of Cleaner Production,2021,308:127287．

[60]LIBARDI N,SOCCOL C R,de CARVALHO J C,et al.Simultaneous cellulase production using domestic wastewater and bioprocess effluent treatment-a biorefinery approach[J].Bioresource Technology,2019,276:42-50．

[61]PAMANIK S K,MAHMUD S,PAUL G K,et al.Fermentation optimization of cellulase production from sugarcane bagasse by Bacillus pseudomycoides and molecular modeling study of cellulase[J].Current Research in Microbial Sciences,2021,2:100013．

[62]McMEEKIN T A,OLLEY J,RATKOWSKY D A,et al.Predictive microbiology:towards the interface and beyond[J].International Journal of Food Microbiology,2002,73:395-407．

[63]陈燕飞．pH对微生物的影响[J]．太原师范学院学报（自然科学版），2009,8(3):121-124,131.CHEN Y F.pH to uygur biology influence[J].Journal of Taiyuan Normal University(Natural Science Edition),2009,8(3):121-124,131．

[64]刘军平．聚乙二醇修饰对去折叠胰蛋白酶的影响[D]．南昌：南昌大学，2012.LIU J P.Effect of PEG modification on unfolded trypsin[D].Nanchang:Nanchang University,2012．

[65]余圆圆．纤维素酶修饰及其对纤维素纤维作用的研究[D]．无锡：江南大学，2014.YU Y Y.Modification of cellulase and its action on cellulose fiber[D].Wuxi:Jiangnan University,2014．

[66]PARK J W,PARK K,SONG H,et al.Saccharification and adsorption characteristics of modified cellulases with hydrophilic/hydrophobic copolymers[J].Journal of Biotechnology,2002,93:203-208．

[67]张裕卿，李滨县．聚氧乙烯-马来酸酐聚合物改性纤维素酶催化酸水解盾叶薯蓣薯蓣皂苷元的研究[J]．中草药，2007,38(4):527-530.ZHANG Y Q,LI B X.Biocatalytic acid hydrolysis of diosgenin in Dioscorea zingiberensis through polyoxethylene oxide-maleic acid anhydride modified cellulose enzyme[J].Chinese Traditional and Herbal Drugs,2007,38(4):527-530．

[68]BUND R K,SINGHAL R S.An alkali stable cellulase by chemical modification using maleic anhydride[J].Carbohydrate Polymers,2002,47(2):137-141．

[69]ZHENG Y J,WANG X Y,SUN Y,et al.Effects of ultrafine grinding and cellulase hydrolysis separately combined with hydroxypropylation,carboxymethylation and phosphate crosslinking on the in vitro hypoglycaemic and hypolipidaemic properties of millet bran dietary fibre[J].LWT,2022,172:114210．

[70]LI T T,GONG X W,YANG G S,et al.Cross-linked enzyme aggregates(CLEAs)of cellulase with improved catalytic activity,adaptability and reusability[J].Bioprocess and Biosystems Engineering,2022,45(5):865-875．

[71]YANG Y,GHATGE S,HUR H G.Enhanced enzyme activity of thermoalkaliphilic laccase(CtLac)of Caldalkalibacillus thermarumstrain TA2.A1by random and site-directed mutagenesis and its application to lignin degradation[J].2022.DOI:https：//doi.org/10.21203/rs.3.rs-1993592/v1．

[72]ZHOU H Y,YI X N,CHEN Q,et al.Improvement of catalytic performance of endoglucanase CgEndo fromColletotrichum graminicola by site-directed mutagenesis[J].Enzyme and Microbial Technology,2022,154:109963．

[73]PAZARLIOGˇLU N K,SARII?IK M,TELEFONCU A.Treating denim fabrics with immobilized commercial cellulases[J].Process Biochemistry,2005,40(2):767-771．

[74]LEE C K,DARAH I,IBRAHIM C O.Enzymatic deinking of laser printed office waste papers:some governing parameters on deinking efficiency[J].Bioresource Technology,2007,98(8):1684-1689．

[75]何芳芳，王海军，王雪莹．纤维素酶的研究进展[J]．造纸科学与技术，2020,39(4):1-8.HE F F,WANG H J,WANG X Y.Research progress of cellulase[J].Paper Science and Technology,2020,39(4):1-8．

[76]LYU D,WANG W,WEI D Z.Construction of two vectors for gene expression in Trichoderma reesei[J].Plasmid,2012,67(1):67-71．

[77]HSU J C,LAKHANI N N.Softer and higher strength paper products and methods of making such products:US 20020162635[P].2002-11-07．

[78]DOERNER K C,WHITE B A.Assessment of the endo-1,4-β-glucanase components of Ruminococcus flavefaciens FD-1[J].Applied and Environmental Microbiology,1990,56(6):1844-1850．

[79]王全军，陈勇，朱伟云．功能性低聚糖对动物机体的免疫作用[J]．粮食与饲料工业，2000(6):35-36.WANG Q J,CHEN Y,ZHU W Y.The immune effect of functional oligosaccharides on animal organism[J].Cereal&Feed Industry,2000(6):35-36．

[80]GIRALDO L A,TEJIDO M L,RANILLA M J,et al.Effects of exogenous cellulase supplementation on microbial growth and ruminal fermentation of a high-forage diet in Rusitec fermenters[J].Journal of Animal Science,2007,85(8):1962-1970．

[81]白晰．圆唇散白蚁肠道微生物内切β-葡聚糖酶工程菌的构建及其酶学性质研究[D]．南京：南京农业大学，2017.BAI X.Expression and characterization of an endo-glucanase from symbiotic bacteria of Reticulitermes labralis[D].Nanjing:Nanjing Agricultural University,2017．

[82]BRENES A,SLOMINSKI B A,MARQUARDT R R,et al.Effect of enzyme addition on the digestibilities of cell wall polysaccharides and oligosaccharides from whole,dehulled,and ethanol-extracted white lupins in chickens[J].Poultry Science,2003,82(11):1716-1725．

[83]BAMFORTH C W.Current perspectives on the role of enzymes in brewing[J].Journal of Cereal Science,2009,50(3):353-357．

[84]姜淑荣．浅谈纤维素酶在酒类生产中的应用[J]．中国酿造，2008(5):12-15.JIANG S R.Brief review on the application of cellulase in the production of liquors[J].China Brewing,2008(5):12-15．

[85]KUHAD R C,GUPTA R,SINGH A.Microbial cellulases and their industrial applications[J].Enzyme Research,2011,2011:280696．

[86]ANGENENT L T,KARIM K,AL-DAHHAN M H,et al.Production of bioenergy and biochemicals from industrial and agricultural wastewater[J].Trends in Biotechnology,2004,22(9):477-485．

[87]ZHANG Y H P,HIMMEL M E,MIELENZ J R.Outlook for cellulase improvement:screening and selection strategies[J].Biotechnology Advances,2006,24(5):452-481.